Nick Garbutt / Male O'Shaughnessy's Chameleon (Calumma oshaughnessyi) at night. Vohiparara, Ranomafana National Park, south eastern Madagascar, November.

Alan Watson / Female O'Shaughnessy's chameleon (Calumma oshaughnessyi) on tree branch at 1,200 metres, Ranomafana National Park, Madagascar

Thomas Marent / O'shaughnessy's Chameleon (Calumma oshaughnessyi) female, Montagne D'Ambre National Park, Antsiranana, northern Madagascar

Thomas Marent / O'shaughnessy's Chameleon (Calumma oshaughnessyi) male at night, Montagne D'Ambre National Park, Antsiranana, northern Madagascar

Jouan & Rius / O'Shaughnessy's Chameleon (Calumma oshaughnessy) at night, tropical rainforest, Montagne d'Ambre NP, Madagascar

Pete Oxford / O'shaughnessy's chameleon (Calumma oshaughnessyi) night time sleeping position, Andohahela National Park, MADAGASCAR, endemic

Pete Oxford / O'shaughnessy's chameleon (Calumma oshaughnessyi) male in breeding colours, Andohahela National Park, MADAGASCAR, endemic

Pete Oxford / O'shaughnessy's chameleon (Calumma oshaughnessyi) male in bright breeding colours, Andohahela National Park, MADAGASCAR, endemic

Pete Oxford / O'shaughnessy's chameleon (Calumma oshaughnessyi) male in breeding colours, Andohahela National Park, MADAGASCAR, endemic

Pete Oxford / O'shaughnessy's chameleon (Calumma oshaughnessyi) male in bright breeding colours, Andohahela National Park, MADAGASCAR, endemic

Pete Oxford / O'shaughnessy's chameleon (Calumma oshaughnessyi) males fighting, Andohahela National Park, MADAGASCAR, endemic

Nick Garbutt / O'Shaughnessy's chameleon (Calumma oshaughnessysi dabrensis). Madagascar

Nick Garbutt / O'Shaughnessy's chameleon (Calumna oshaughnessyi), Ranomafana NP, Madagascar